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Note the great degree of individual variation in fur colour, which was once used as a basis for separating the species into various subspecies. Unlike the striped hyena, for which a number of subspecies were proposed in light of its extensive modern range, the spotted hyena is a genuinely variable species, both temporally and spatially. Its range once encompassed almost all of Africa and Eurasia , and displayed a large degree of morphological geographic variation, which led to an equally extensive set of specific and subspecific epithets. It was gradually realised that all of this variation could be applied to individual differences in a single subspecies. In , biologist L. Harrison Matthews demonstrated through comparisons between a large selection of spotted hyena skulls from Tanzania that all the variation seen in the then recognised subspecies could also be found in a single population, with the only set of characters standing out being pelage which is subject to a high degree of individual variation and size which is subject to Bergmann's Rule.
Pinterest Dr Charles Runels center and his team. Bryan Tarnowski for the Guardian As we awaited the arrival of my first interview subject, Lacey, Runels suggested that I might want to try the O-Shot for myself. She told me that doctors regularly flew in from all over the world to be trained in the procedure, and that initially they reacted to offers of free O-Shots just as I had — with a mix of embarrassment and surprise. But by the end, she said, everyone wanted one. She and Runels laughed, recalling how the last class had run until three in the morning, just to accommodate demand.